juice, which in these animals is unusually copious. Thus softened and rendered tender, they are returned a second time to the action of the mouth, where the grinding teeth complete at their leisure the trituration which is necessary, but which was before left imperfect. I the trituration which is necessay, sary; for it appears from experiments, that the gastric fluid of sheep, for example, has no effect in digesting plants, unless they have been previously masticated; that it only produces a slight maceration, nearly as common water would do in a like degree of heat ; but that when once vegetables are reduced to pieces by mastication, the fluid then exerts upon them its specific operation. Its first effect is to soften them, and to destroy their natural consistency; it then goes on to dissolve them; not sparing even the toughest parts, such as the nerves of the leaves*.

I think it very probable, that the gratification also of the animal is renewed and prolonged by this faculty. Sheep, deer, and oxen, appear to be in a state of enjoyment whilst they are chewing the cud. It is then, perhaps, that they best relish their food.

II. In birds, the compensation is still more striking. They have no teeth at all. What *Spall. Dis. iii. sect. cxl.

have they then to make up for this severe want? I speak of granivorous and herbivorous birds; such as common fowls, turkeys, ducks, geese, pigeons, &c.; for it is concerning these alone that the question need be asked. All these are furnished with a peculiar and most powerful muscle, called the gizzard; the inner coat of which is fitted up with rough plaits, which, by a strong friction against one another, break and grind the hard aliment as effectually, and by the same mechanical action, as a coffee-mill would do. It has been proved by the most correct experiments, that the gastric juice of these brids will not operate upon the entire grain; not even when softened by water or macerated in the crop. Therefore without a grinding-machine within its body, without the trituration of the gizzard, a chicken would have starved upon a heap of corn. Yet why should a bill and a gizzard go together? Why should a gizzard never be found where there are teeth?

Nor does the gizzard belong to birds as such. A gizzard is not found in birds of prey. Their food requires not to be ground down in a mill. The compensatory contrivance goes no further than the necessity. In both classes of birds, however, the digestive organ within the body bears a strict and

mechanical relation to the external instruments for procuring food. The soft membranous stomach accompanies a hooked, notched beak; short, muscular legs; strong, sharp, crooked talons: the cartilaginous stomach attends that conformation of bill and toes, which restrains the bird to the picking of seeds, or the cropping of plants.

III. But to proceed with our compensations. A very numerous and comprehensive tribe of terrestrial animals are entirely without feet; yet locomotive; and in a very considerable degree swift in their motion. How is the want of feet compensated? It is done by the disposition of the muscles and fibres of the trunk. In consequence of the just collocation, and by means of the joint action of longitudinal and annular fibres, that is to say, of strings and rings, the body and train of reptiles are capable of being reciprocally shortened and lengthened, drawn up and stretched out. The result of this action is a progressive, and, in some cases, a rapid movement of the whole body, in any direction to which the will of the animal determines it. The meanest creature is a collection of wonders. The play of the rings in an earthworm, as it crawls; the undulatory motion propagated along the body; the beards or

U

prickles with which the annuli are armed, and which the animal can either shut up close to its body, or let out to lay hold of the roughness of the surface upon which it creeps; and the power arising from all these, of changing its place and position, afford, when compared with the provisions for motion in other animals, proofs of new and appropriate mechanism. Suppose that we had never seen an animal move upon the ground without feet, and that the problem was; muscular action, i. e. reciprocal contraction and relaxation being given, to describe how such an animal might be constructed, capable of voluntarily changing place. Something, perhaps, like the organization of reptiles, might have been hit upon by the ingenuity of an artist; or might have been exhibited in an automaton, by the combination of springs, spiral wires, and ringlets: but to the solution of the problem would not be denied, surely, the praise of invention and of successful thought: least of all could it ever be questioned, whether intelligence had been employed about it, or not.

CHAPTER XVII.

THE RELATION OF ANIMATED BODIES TO INANIMATE

NATURE.

WE have already considered relation, and under different views; but it was the relation

of parts to parts, of the parts of an animal to other parts of the same animal, or of another individual of the same species.

But the bodies of animals hold, in their constitution and properties, a close and important relation to natures altogether external to their own; to inanimate substances, and to the specific qualities of these; e. g. they hold a strict relation to the ELEMENTS by which they are surrounded.

I. Can it be doubted, whether the wings of birds bear a relation to air, and the fins of fish to water? They are instruments of motion, severally suited to the properties of the medium in which the motion is to be performed: which properties are different. Was not this difference contemplated, when the instruments were differently constituted?

II. The structure of the animal ear depends for its use not simply upon being surrounded