fly belonging to a fourth genus. It deserves especial notice that many of the mimicking forms of the Leptalis, as well as of the mimicked forms, can be shown by a graduated series to be merely varieties of the same. species; whilst others are undoubtedly distinct species. But why, it may be asked, are certain forms treated as the mimicked and others as the mimickers? Mr. Bates satisfactorily answers this question, by showing that the form which is imitated keeps the usual dress of the group to which it belongs, whilst the counterfeiters have changed their dress and do not resemble their nearest allies.

We are next led to inquire what reason can be assigned for certain butterflies and moths so often assuming the dress of another and quite distinct form; why, to the perplexity of naturalists, has nature condescended. to the tricks of the stage? Mr. Bates has, no doubt, hit on the true explanation. The mocked forms, which always abound in numbers, must habitually escape destruction to a large extent, otherwise they could not exist in such swarms; and a large amount of evidence has now been collected, showing that they are distasteful to birds and other insect-devouring animals. The mocking forms, on the other hand, that inhabit the same district, are comparatively rare, and belong to rare groups; hence they must suffer habitually from some danger, for otherwise, from the number of eggs laid by all butterflies, they would in three or four generations swarm over the whole country. Now if a member of one of these persecuted and rare groups were to assume a dress so like that of a well-protected species that it continually deceived the practised eyes of an entomologist, it would often deceive predaceous birds

and insects, and thus often escape destruction. Mr. Bates may almost be said to have actually witnessed the process by which the mimickers have come so closely to resemble the mimicked; for he found that some of the forms of Leptalis which mimic so many other butterflies, varied in an extreme degree. In one district several varieties occurred, and of these one alone resembled to a certain extent, the common Ithomia of the same district. In another district there were two or three varieties, one of which was much commoner than the others, and this closely mocked another form of Ithomia. From facts of this nature, Mr. Bates concludes that the Leptalis first varies; and when a variety happens to resemble in some degree any common butterfly inhabiting the same district, this variety, from its resemblance to a flourishing and little-persecuted kind, has a better chance of escaping destruction from predaceous birds and insects, and is consequently oftener preserved;" the less perfect degrees of resemblance being generation after generation eliminated, and only the others left to propagate their kind." that here we have an excellent illustration of natural selection.

Messrs. Wallace and Trimen have likewise described several equally striking cases of imitation in the Lepidoptera of the Malay Archipelago and Africa, and with some other insects. Mr. Wallace has also detected one such case with birds, but we have none with the larger quadrupeds. The much greater frequency of imitation with insects than with other animals, is probably the consequence of their small size; insects cannot defend themselves, excepting indeed the kinds furnished with a string, and I have never heard of an instance of

such kinds mocking other insects, though they are mocked; insects cannot easily escape by flight from the larger animals which prey on them; therefore, speaking metaphorically, they are reduced, like most weak creatures, to trickery and dissimulation.

It should be observed that the process of imitation probably never commenced between forms widely dissimilar in colour. But starting with species already somewhat like each other, the closest resemblance, if beneficial, could readily be gained by the above means; and if the imitated form was subsequently and gradually modified through any agency, the imitating form would be led along the same track, and thus be altered to almost any extent, so that it might ultimately assume an appearance or colouring wholly unlike that of the other members of the family to which it belonged. There is, however, some difficulty on this head, for it is necessary to suppose in some cases that ancient members belonging to several distinct groups, before they had diverged to their present extent, accidentally resembled a member of another and protected group in a sufficient degree to afford some slight protection; this having given the basis for the subsequent acquisition of the most perfect resemblance.

On the Nature of the Affinities connecting Organic Beings. As the modified descendants of dominant species, belonging to the larger genera, tend to inherit the advantages which made the groups to which they belong large and their parents dominant, they are almost sure to spread widely, and to seize on more and more places in the economy of nature. The larger and more dominant groups within each class thus tend to

go on increasing in size; and they consequently supplant many smaller and feebler groups. Thus we can account for the fact that all organisms, recent and extinct, are included under a few great orders, and under still fewer classes. As showing how few the higher groups are in number, and how widely they are spread throughout the world, the fact is striking that the discovery of Australia has not added an insect belonging to a new class; and that in the vegetable kingdom, as I learn from Dr. Hooker, it has added only two or three families of small size.

In the chapter on Geological Succession I attempted to show, on the principle of each group having generally diverged much in character during the long-continued process of modification, how it is that the more ancient forms of life often present characters in some degree intermediate between existing groups. As some few of the old and intermediate forms have transmitted to the present day descendants but little modified, these constitute our so-called osculant or aberrant species. The more aberrant any form is, the greater must be the number of connecting forms which have been exterminated and utterly lost. And we have some evidence of aberrant groups having suffered severely from extinction, for they are almost always represented by extremely few species; and such species as do occur are generally very distinct from each other, which again. implies extinction. The genera Ornithorhynchus and Lepidosiren, for example, would not have been less aberrant had each been represented by a dozen species, instead of as at present by a single one, or by two or three. We can, I think, account for this fact only by looking at aberrant groups as forms which have been con

quered by more successful competitors, with a few members still preserved under unusually favourable conditions.

Mr. Waterhouse has remarked that, when a member belonging to one group of animals exhibits an affinity to a quite distinct group, this affinity in most cases is general and not special; thus, according to Mr. Waterhouse, of all Rodents, the bizcacha is most nearly related to Marsupials; but in the points in which it approaches this order, its relations are general, that is, not to any one marsupial species more than to another. As these points of affinity are believed to be real and not merely adaptive, they must be due in accordance with our view to inheritance from a common progenitor. Therefore we must suppose either that all Rodents, including the bizcacha, branched off from some ancient Marsupial, which will naturally have been more or less intermediate in character with respect to all existing Marsupials; or that both Rodents and Marsupials branched off from a common progenitor, and that both groups have since undergone much modification in divergent directions. On either view we must suppose that the bizcacha has retained, by inheritance, more of the characters of its ancient progenitor than have other Rodents; and therefore it will not be specially related to any one existing Marsupial, but indirectly to all or nearly all Marsupials, from having partially retained the character of their common progenitor, or of some early member of the` group. On the other hand, of all Marsupials, as Mr. Waterhouse has remarked, the Phascolomys resembles most nearly, not any one species, but the general order of Rodents. In this case, however, it may be strongly suspected that the resemblance is only analogical, owing