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mosomes are found associated in pairs distributed over a network. The further history of the reduction mitoses is not described, so that the investigation is incomplete in a number of important features.

The second paper of Rosenberg1 deals with Crepis virens, one of the Composita, a form remarkable for the small number of chromosomes, which are six for the sporophyte and three for the gametophyte generation. A further important peculiarity is a difference in the size of the chromosomes which makes it possible to follow the individual elements through succeeding mitoses with some degree of certainty. This is, so far as the reviewer is aware, the first account for plants of such a differentiation of chromosomes as has been described for animals by a number of zoologists.

The nuclei of the sporophyte (somatic) show six small prochromosomes in the resting stage from which are organized during the prophases of the vegetative mitoses two short rod-shaped chromosomes, two very long bent elements, and two chromosomes about midway in length between these extremes. The resting nuclei of the pollen mother-cells have six prochromosomes more or less clearly grouped in pairs. Synapsis presents a series of parallel threads intimately united at intervals. From this condition a thick coiled spirem is organized which clearly shows its double nature in the frequent longitudinal separation of portions as though it were split. The free ends of the chromosomes composing the spirem may at times be distinguished. A gradual contraction of the spirem leads through stages comparable to those described as a second contraction by various authors to the period when the six chromosomes, grouped in three pairs, may be clearly recognized (diakinesis).

The chromosome group on the approach of the heterotypic mitosis consists then of a pair of small, almost spherical, chromosomes, a pair of long rods, and a pair of short rods. These correspond to the three different sizes of chromosomes present in the vegetative sporophytic mitoses, but are more condensed or shortened. Thus the heterotypic mitosis is a true reduction division distributing the six chromosomes in two sets each of which consists of a spherical chromosome, a long rod, and a chromosome intermediate in shape between these two. These

Rosenberg, O., "Zur Kenntniss von den Tetradenteilungen der Compositen," Svensk Bot. Tidsk., III, p. 64, 1909.

chromosomes divide during the anaphase of the heterotypic mitosis in preparation for the second or homotypic mitosis so that they appear at the poles of the heterotypic spindle in the form of three split chromosomes or pairs.

The chromosomes change their form in the interkinesis, becoming long spiral threads which shorten on the approach of the homotypic mitosis when the six chromosomes again appear as three pairs showing the same characteristic range of form. The members of these three pairs are distributed by this division so that the nucleus of each pollen grain receives three chromosomes, a short, a long, and a middle-sized element, and these may be recognized in the resting nucleus by three prochromosomes. A brief examination of the mitoses in the embryo-sac supported the conclusions above outlined.

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